Below are lists of extant fern families and subfamilies using the classification scheme proposed by the Pteridophyte Phylogeny Group in 2016 (PPG I).[1] The scheme is based on molecular phylogenetic studies, and also draws on earlier classifications,[1] particularly those by Smith et al. (2006),[2] Chase and Reveal (2009),[3] and Christenhusz et al. (2011).[4] It rejects the very broad circumscription of some families used by Christenhusz and Chase (2014), whose Aspleniaceae corresponds to the entire suborder Aspleniineae of PPG I.[1][5]
Systematic order
[edit]- Class Polypodiopsida Cronquist, Takht. & W.Zimm. (ferns, including horsetails)
- Subclass Equisetidae Warm. (horsetails and scouring-rushes)
- Order Equisetales DC. ex Bercht. & J.Presl
- Family Equisetaceae Michx. ex DC
- Subclass Ophioglossidae Klinge (whisk ferns, adder's-tongues and moonworts)
- Order Psilotales Prant
- Family Psilotaceae J.W.Griff. & Henfr.
- Order Ophioglossales Link
- Family Ophioglossaceae Martinov
- Subfamily Helminthostachyoideae C.Presl
- Subfamily Mankyuoideae J.R.Grant & B.Dauphin
- Subfamily Ophioglossoideae C.Presl
- Subfamily Botrychioideae C.Presl
- Subclass Marattiidae Klinge (marattioid ferns)
- Order Marattiales Link
- Family Marattiaceae Kaulf
- Subclass Polypodiidae Cronquist, Takht. & W.Zimm. (leptosporangiate ferns)
- Order Osmundales Link (royal ferns)
- Family Osmundaceae Martinov
- Order Hymenophyllales A.B.Frank (filmy ferns and bristle ferns)
- Family Hymenophyllaceae Mart
- Subfamily Trichomanoideae C.Presl
- Subfamily Hymenophylloideae Burnett
- Order Gleicheniales Schimp
- Family Matoniaceae C.Pres
- Family Dipteridaceae Seward & E.Dale
- Family Gleicheniaceae C.Presl
- Order Schizaeales Schimp.
- Family Lygodiaceae M.Roem
- Family Schizaeaceae Kaulf
- Family Anemiaceae Link
- Order Salviniales Link
- Family Salviniaceae Martinov
- Family Marsileaceae Mirb.
- Order Cyatheales A.B.Frank (tree ferns)
- Family Thyrsopteridaceae C.Presl
- Family Loxsomataceae C.Presl
- Family Culcitaceae Pic.Serm
- Family Plagiogyriaceae Bowe
- Family Cibotiaceae Koral
- Family Metaxyaceae Pic.Serm.
- Family Dicksoniaceae M.R.Schomb.
- Family Cyatheaceae Kaulf.
- Order Polypodiales Link
- Suborder Saccolomatineae Hovenkamp
- Family Saccolomataceae Doweld
- Suborder Lindsaeineae Lehtonen & Tuomist
- Family Cystodiaceae J.R.Croft
- Family Lonchitidaceae Doweld
- Family Lindsaeaceae C.Presl ex M.R.Schomb.
- Suborder Pteridineae J.Prado & Schuettp
- Family Pteridaceae E.D.M.Kirchn.
- Subfamily Parkerioideae Burnett
- Subfamily Cryptogrammoideae S.Lindsay
- Subfamily Pteridoideae Link
- Subfamily Vittarioideae Link
- Subfamily Cheilanthoideae Horvat
- Suborder Dennstaedtiineae Schwartsb. & Hovenkamp
- Family Dennstaedtiaceae Lotsy
- Suborder Aspleniineae H.Schneid. & C.J.Rothf
- Family Cystopteridaceae Shmakov
- Family Rhachidosoraceae X.C.Zhang
- Family Diplaziopsidaceae X.C.Zhang & Christenh.
- Family Desmophlebiaceae Mynssen
- Family Hemidictyaceae Christenh. & H.Schneid.
- Family Aspleniaceae Newman
- Family Woodsiaceae Herter
- Family Onocleaceae Pic.Serm.
- Family Blechnaceae Newman
- Subfamily Stenochlaenoideae (Ching) J.P.Roux
- Subfamily Woodwardioideae Gasper
- Subfamily Blechnoideae Gasper, V.A.O.Dittrich & Salino
- Family Athyriaceae Alston
- Family Thelypteridaceae Ching ex Pic.Serm.
- Subfamily Phegopteridoideae Salino, A.R.Sm. & T.E.Almeid
- Subfamily Thelypteridoideae C.F.Reed
- Suborder Polypodiineae Dumort.
- Family Didymochlaenaceae Ching ex Li Bing Zhang & Liang Zhang
- Family Hypodematiaceae Ching
- Family Dryopteridaceae Herter
- Subfamily Polybotryoideae H.M.Liu & X.C.Zhang
- Subfamily Elaphoglossoideae (Pic.Serm.) Crabbe, Jermy & Mickel
- Subfamily Dryopteridoideae Link
- 2 genera unassigned to a subfamily
- Family Nephrolepidaceae Pic.Serm.
- Family Lomariopsidaceae Alston
- Family Tectariaceae Panigrahi
- Family Oleandraceae Ching ex Pic.Serm.
- Family Davalliaceae M.R.Schomb.
- Family Polypodiaceae J.Presl & C.Presl
- Subfamily Loxogrammoideae H.Schneid.
- Subfamily Platycerioideae B.K.Nayar
- Subfamily Drynarioideae Crabbe, Jermy & Mickel
- Subfamily Microsoroideae B.K.Nayar
- Subfamily Polypodioideae Sweet
- Subfamily Grammitidoideae Parris & Sundue
- 1 genus unassigned to a subfamily
Alphabetic order
[edit]- Anemiaceae Link (1 genus)
- Aspleniaceae Newman (2 genera)
- Athyriaceae Alston (3 genera)
- Blechnaceae Newman (24 genera)
- Cibotiaceae Koral (1 genus)
- Culcitaceae Pic.Serm (1 genus)
- Cyatheaceae Kaulf. (3 genera)
- Cystodiaceae J.R.Croft (1 genus)
- Cystopteridaceae Shmakov (3 genera)
- Davalliaceae M.R.Schomb. (1 genus)
- Dennstaedtiaceae Lotsy (10 genera)
- Desmophlebiaceae Mynssen (1 genus)
- Dicksoniaceae M.R.Schomb. (3 genera)
- Didymochlaenaceae Ching ex Li Bing Zhang & Liang Zhang (1 genus)
- Diplaziopsidaceae X.C.Zhang & Christenh. (2 genera)
- Dipteridaceae Seward & E.Dale (2 genera)
- Dryopteridaceae Herter (26 genera)
- Equisetaceae Michx. ex DC (1 genus)
- Gleicheniaceae C.Presl (6 genera)
- Hemidictyaceae Christenh. & H.Schneid. (1 genus)
- Hymenophyllaceae Mart (9 genera)
- Hypodematiaceae Ching (2 genera)
- Lindsaeaceae C.Presl ex M.R.Schomb. (7 genera)
- Lomariopsidaceae Alston (4 genera)
- Lonchitidaceae Doweld (1 genus)
- Loxsomataceae C.Presl (2 genera)
- Lygodiaceae M.Roem (1 genus)
- Marattiaceae Kaulf (6 genera)
- Marsileaceae Mirb. (3 genera)
- Matoniaceae C.Pres (2 genera)
- Metaxyaceae Pic.Serm. (1 genus)
- Nephrolepidaceae Pic.Serm. (1 genus)
- Oleandraceae Ching ex Pic.Serm. (1 genus)
- Onocleaceae Pic.Serm. (4 genera)
- Ophioglossaceae Martinov (10 genera)
- Osmundaceae Martinov (6 genera)
- Plagiogyriaceae Bowe (1 genus)
- Polypodiaceae J.Presl & C.Presl (65 genera)
- Psilotaceae J.W.Griff. & Henfr. (2 genera)
- Pteridaceae E.D.M.Kirchn. (53 genera)
- Rhachidosoraceae X.C.Zhang (1 genus)
- Saccolomataceae Doweld (1 genus)
- Salviniaceae Martinov (2 genera)
- Schizaeaceae Kaulf (2 genera)
- Tectariaceae Panigrahi (7 genera)
- Thelypteridaceae Ching ex Pic.Serm. (30 genera)
- Thyrsopteridaceae C.Presl (1 genus)
- Woodsiaceae Herter (1 genus)
See also
[edit]References
[edit]- ^ a b c PPG I (2016). "A community-derived classification for extant lycophytes and ferns". Journal of Systematics and Evolution. 54 (6): 563–603. doi:10.1111/jse.12229.
- ^ Alan R. Smith; Kathleen M. Pryer; Eric Schuettpelz; Petra Korall; Harald Schneider; Paul G. Wolf (2006). "A classification for extant ferns" (PDF). Taxon. 55 (3): 705–731. doi:10.2307/25065646. Archived from the original (PDF) on 2008-02-26.
- ^ Mark W. Chase & James L. Reveal (2009). "A phylogenetic classification of the land plants to accompany APG III". Botanical Journal of the Linnean Society. 161: 122–127. doi:10.1111/j.1095-8339.2009.01002.x.
- ^ Maarten J. M. Christenhusz, Xian-Chun Zhang & Harald Schneider (2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19: 7–54.
- ^ Christenhusz, Maarten J.M. & Chase, Mark W. (2014). "Trends and concepts in fern classification". Annals of Botany. 113 (9): 571–594. doi:10.1093/aob/mct299. PMC 3936591. PMID 24532607.