Fossil localities in Mongolia. From C to D, mainly Bayan Shireh locations
The Bayan Shireh Formation (also known as Baynshiree/Baynshire, Baynshirenskaya Svita or Baysheen Shireh) is a geological formation in Mongolia, that dates to the Cretaceous period. It was first described and established by Vasiliev et al. 1959.[1]
The Bayan Shireh Formation is primarily composed by varicoloured claystones and sandstones with calcareous concretions and characterized by grey mudstones and yellowish-brown medium grained sandstones.[1][2] Up to 300 m (980 ft) thick, the most complete sections are found in the eastern Gobi Desert, consisting of fine-grained, often cross-stratified gray sandstone interbedded with claystone and concretionary, intraformational conglomerates with relatively thick units of red to brown mudstone in the upper part.[3][4] The Baynshire and Burkhant localities are mainly composed by mudstone, siltstone, sandstone, and conglomerates, with most of their sedimentation being fluvial.[5][6][7] The environments that were present on the Bayan Shireh Formation consisted mainly on semi-arid climates with large water bodies, such as rivers or lakes.[8][7] Although it is considered to be partially lacustrine,[9] largescale cross-stratification in many of the sandstone layers at the Baynshire locality seem to indicate that a large meanderingfluvial system was present. It has been implied that during the late Bayan Shirehnian times, large rivers with direct connections to the sea drained a prominent part of the eastern Gobi region.[3][6][8]
According to Jerzykiewicz and Russell, the Bayan Shireh Formation can be divided into 2 informal units: upper and lower beds.[5] Danilov and colleagues have suggested that the lower beds are Cenomanian to early Turonian, and the upper beds are late Turonian to Santonian in age.[10] While the lower beds are composed by extensive conglomerate that indicates the ancient presence of very active rivers, the upper beds are mainly composed by mudstone and claystone that is interbedded by sandstone, indicating again, the presence of rivers but also lakes and other water bodies.[11]
Based on comparisons with other formations, Jerzykiewicz and Russell suggested that the Bayan Shireh paleofauna seems to correspond best with the Turonian through early Campanian stages of the Late Cretaceous, about 93 to 80 million years ago.[5] However, examination of the magnetostratigraphy of the formation indicates that the entire Bayan Shireh lies within the Cretaceous Long Normal, which lasted only until the end of the Santonian stage, giving a possible Cenomanian through Late Santonian age, between 98 and 83 million years ago.[6] In 2012, Averianov and Sues re-examined many formations from the Gobi Desert and using biostratigraphic occurrences and previous dating, the Bayan Shireh Formation was considered to be Cenomanian to Santonian in age. The lower beds dating to 98 million and 90 million years ago (early Cenomanian to late Turonian), and the upper beds dating to 90 million and 83 million years ago (late Turonian to late Santonian).[8]CalciteU–Pb measurements performed by Kurumada and colleagues in 2020 have estimated the exact age of the Bayan Shireh Formation from 102–86 Ma, based on dates obtained of 95.9 ± 6.0 Ma and 89.6 ± 4.0 Ma.[12]
A potential correlation between the Iren Dabasu Formation has been long suggested by most authors, mainly based on the highly similar vertebrate assemblages.[5][13][14][15] However, Van Itterbeeck et al. 2005 argued against this correlation concluding that instead, the Iren Dabasu Formation was coeval with the younger Nemegt Formation based on the charophyte and ostracode assemblages; therefore, these formations were dated to the Late Campanian-Early Maastrichtian.[16] Averianov and Sues instead proposed a correlation between the Bayan Shireh, Iren Dabasu and Bissekty formations.[8] In 2015, Tsuihiji and team found the Iren Dabasu-Nemegt correlation to be inconsistent since the microfossil assemblages used by Van Itterbeeck and colleagues were not restricted to the Maastrichtian period and the similarities between these assemblages were most probably due to a similar deposition and climate settings.[17]
In terms of biodiversity, therizinosaurs and turtles were the most abundant vertebrates across the formation, as evidenced on numerous remains.[10][18] Nevertheless, hadrosauroids were fairly abundant too, particularly at the Baynshire locality with numerous remains unearthed from this area and a new unnamed hadrosauroid. In addition, most specimens of Gobihadros come from this locality.[7][19][11] Also, niche partitioning has been reported within the therizinosauridsErlikosaurus and Segnosaurus,[20] and the ankylosaursTalarurus and Tsagantegia.[21]Mammals however, are extremely uncommon; Tsagandelta is the only mammal described so far.[22] Besides vertebrate fossils, abundant fossil fruits have been collected from the Bor Guvé and Khara Khutul localities and they are especially abundant at Bor Guvé. Although they resemble Abelmoschus esculentus their taxonomic position remains unclear and further examination is required.[23]
^ abVasiliev, V. G.; Volkhonin, V. C.; Grishin, G. L.; Ivanov, A. Kh.; Marinov, I. A.; Mokshancev, K. B. (1959). Geological structure of the People's Republic of Mongolia (stratigraphy and tectonics). p. 492.
^Watabe, M.; Suzuki, S. (2000). "Report on the Japan - Mongolia Joint Paleontological Expedition to the Gobi desert, 1994". Hayashibara Museum of Natural Sciences, Research Bulletin. 1: 30−44.
^ abSochava, A. V. (1975). "Stratigraphy and lithology of the Upper Cretaceous sediments in southern Mongolia. In Stratigraphy of Mesozoic sediments of Mongolia". Transactions of Joint Soviet–Mongolian Scientific Research and Geological Expedition. 13: 113–182.
^Martinson, G. G. (1982). "The Upper Cretaceous mollusks of Mongolia". Sovmestnaya Sovetsko– Mongolskaya Paleontolog-icheskaya Ekspeditsia. 17: 5–76.
^ abcdDanilov, I. G.; Hirayama, R.; Sukhanov, V. B.; Suzuki, S.; Watabe, M.; Vitek, N. S. (2014). "Cretaceous soft-shelled turtles (Trionychidae) of Mongolia: new diversity, records, and a revision". Journal of Systematic Palaeontology. 12 (7): 799–832. Bibcode:2014JSPal..12..799D. doi:10.1080/14772019.2013.847870. S2CID86304259.
^ abcdSlowiak, J.; Szczygielski, T.; Ginter, M.; Fostowicz-Frelik, L. (2020). "Uninterrupted growth in a non-polar hadrosaur explains the gigantism among duck-billed dinosaurs". Palaeontology. 63 (4): 579–599. Bibcode:2020Palgy..63..579S. doi:10.1111/pala.12473. S2CID213247742.
^Currie, P. J.; Eberth, D. A. (1993). "Palaeontology, sedimentology and palaeoecology of the Iren Dabasu Formation (Upper Cretaceous), Inner Mongolia, People's Republic of China". Cretaceous Research. 14 (2): 127−144. Bibcode:1993CrRes..14..127C. doi:10.1006/cres.1993.1011.
^Nessov, L. A. (1995). Dinosaurs of northern Eurasia: new data about assemblages, ecology and paleobiogeography (in Russian). Saint Petersburg: Saint Petersburg University. p. 156. Translated paper
^ abLee, Y. N.; Lee, H. J.; Kobayashi, Y.; Carabajal, A. P.; Barsbold, R.; Fiorillo, A. R.; Tsogtbaatar, K. (2019). "Unusual locomotion behaviour preserved within a crocodyliform trackway from the Upper Cretaceous Bayanshiree Formation of Mongolia and its palaeobiological implications". Palaeogeography, Palaeoclimatology, Palaeoecology. 533 (109353): 2. Bibcode:2019PPP...53309239L. doi:10.1016/j.palaeo.2019.109239. S2CID197584839.
^ abcPark, J. Y.; Lee, Y. N.; Currie, P. J.; Kobayashi, Y.; Koppelhus, E.; Barsbold, R.; Mateus, O.; Lee, S.; Kim, S. H. (2019). "Additional skulls of Talarurus plicatospineus (Dinosauria: Ankylosauridae) and implications for paleobiogeography and paleoecology of armored dinosaurs". Cretaceous Research. 108: 104340. doi:10.1016/j.cretres.2019.104340. S2CID212423361.
^ abRougier, G. W.; Davis, B. M.; Novacek, M. J. (2015). "A deltatheroidan mammal from the Upper Cretaceous Baynshiree Formation, eastern Mongolia". Cretaceous Research. 52: 167–177. Bibcode:2015CrRes..52..167R. doi:10.1016/j.cretres.2014.09.009.
^Efimov, M. B. (1981). "New paralligatorids from the Upper Cretaceous of Mongolia". Trudy Sovmestnoi Sovetsko-Mongol'skoi Paleontologicheskoi Ekspeditsii. 15: 26–28.
^ abKhand, Y.; Badamgarav, D.; Ariunchimeg, Y.; Barsbold, R. (2000). "Cretaceous system in Mongolia and its depositional environments". Cretaceous Environments of Asia. Developments in Palaeontology and Stratigraphy. Vol. 17. pp. 49–79. doi:10.1016/s0920-5446(00)80024-2. ISBN 9780444502766.
^Chkhikvadze, V. M. (1980). "On the question of the origin of the soft−shelled turtles". Bulletin of the Academy of Sciences of the Georgian SSR. 100 (2): 501–503.
^Nessov, L. A. (1981). "Herpetological Investigations in Siberia and the Far East". On the turtle of the family Dermatemydidae from the Cretaceous of Amur River Basin and some other rare findings of remains of ancient turtles of Asia. pp. 69–73.
^Parish, J. C.; Barrett, P. M. (2004). "A reappraisal of the ornithischian dinosaur Amtosaurus magnus Kurzanov and Tumanova 1978, with comments on the status of A. archibaldi Averianov 2002". Canadian Journal of Earth Sciences. 41 (3): 299–306. Bibcode:2004CaJES..41..299P. doi:10.1139/e03-101.
^Maleev, E. A. (1952). "New ankylosaur of the Upper Cretaceous of Mongolia". Doklady Akademii Nauk SSSR. 87: 273−276.
^Tumanova, T. A. (1987). "The armored dinosaurs of Mongolia". The Joint Soviet Mongolian Paleontological Expedition Transaction. 32: 1–76.
^Tumanova, T. A. (2000). "Armoured dinosaurs from the Cretaceous of Mongolia". The Age of Dinosaurs in Russia and Mongolia. Cambridge University Press. pp. 517–532.
^Sereno, P. C. (2000). "The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia". The Age of Dinosaurs in Russia and Mongolia(PDF). Cambridge University Press. pp. 489–491.
^ abcdefBarsbold, R.; Kobayashi, Y.; Kubota, K. (2007). "New discovery of dinosaur fossils from the Upper Cretaceous Bayanshiree Formation of Mongolia". Journal of Vertebrate Paleontology. 27 (supp. 003): 44A. doi:10.1080/02724634.2007.10010458. S2CID220411226.
^Mikhailov, K.; Sabath, K.; Kurzanov, S. (1996). "Egg and nests from the Cretaceous of Mongolia". In Carpenter, K.; Hirsch, K. F.; Horner, J. R. (eds.). Dinosaur Eggs and Babies. Cambridge University Press. pp. 97−98. ISBN 9780521567237.
^Sochava, A. V. (1969). "Dinosaur eggs from the Upper Cretaceous of the Gobi Desert". Paleontological Journal. 1969 (4): 517−527.
^Averianov, A. O; Lopatin, A. V.; Tsogtbaatar, K. (2022). "Taxonomic attribution of a juvenile hadrosauroid dinosaur from the Upper Cretaceous Bayinshire Formation of Mongolia". Doklady Earth Sciences. 503 (1): 93–96. Bibcode:2022DokES.503...93A. doi:10.1134/S1028334X22030035. S2CID246698076.
^Perle, A.; Norell, M. A.; Clark, J. (1999). "A new maniraptoran Theropod−Achillobator giganticus (Dromaeosauridae)−from the Upper Cretaceous of Burkhant, Mongolia". Contributions from the Geology and Mineralogy Chair, National Museum of Mongolia (101): 1–105. OCLC69865262.
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^Watabe, M.; Tsogtbaatar, K.; Suzuki, S.; Saneyoshi, M. (2010). "Geology of dinosaur-fossil-bearing localities (Jurassic and Cretaceous: Mesozoic) in the Gobi Desert: Results of the HMNS-MPC Joint Paleontological Expedition". Hayashibara Museum of Natural Sciences Research Bulletin. 3: 41−118.
^Kubota, K.; Barsbold, R. (2007). "New dromaeosaurid (Dinosauria Theropoda) from the Upper Cretaceous Bayanshiree Formation of Mongolia". Journal of Vertebrate Paleontology. 27 (suppl. to 3): 102A. doi:10.1080/02724634.2007.10010458. S2CID220411226.
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^Chinzorig, T.; Kobayashi, Y.; Saneyoshi, M.; Tsogtbaatar, K.; Batamkhatan, Z.; Ryuji, T. (2017). "Multitaxic bonebed of two new ornithomimids (Theropoda, Ornithomimosauria) from the Upper Cretaceous Bayanshiree Formnation of southeastern Gobi desert, Mongolia". Journal of Vertebrate Paleontology. Program and Abstracts: 97.
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